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Northwest Pacific, Atlantic Ocean, Mediterranean, Antarctic and Arctic Ocean: Iberian Peninsula and Belize. Tropical and subtropical.
Length at first maturity / Size / Weight / Age
Maturity: Lm ?  range ? - ? cm Max length : 3.0 cm TL male/unsexed; (Ref. 415)
Found in rock crevices, mangrove root mats and mud, along subtidal and intertidal zones in marine and brackish environments (Ref. 415). Known to be an osconformer, which is unaffected by large and repeated changes in salinity, but is known to be affected by increases in water temperature (Ref. 86624). Known as a benthic species (Ref. 85346). Intertidal to 590 m in rock crevices; mangrove roots mats and mud (Ref. 415); 34 to 500 m both in Ionian and North Aegean Sea (Ref. 775). Infaunal, burrows (Ref. 116123). This species is collected from mangrove and Thalassia root mats (Ref. 86591).
Life cycle and mating behavior Maturity | Reproduction | Spawning | Eggs | Fecundity | Larvae